Skids is a theoretician. This is appropriate, as some would say his existence is merely theoretical; everyone forgets who he is or that he was there. But if they got to know him, they'd learn that he likes to daydream, get built into furniture, and pick up cowboy chicks.
In the event that the Ark's mission would require more manpower, Optimus Prime had several Autobots encode copies of their personalities as light patterns within crystals, including Skids. When a large number of Prime's troops were then injured during the opening volleys of the war on Earth, Prime arranged for the construction of new bodies, which the personalities were then programmed into, bringing Skids and the others to life "for a second time." Now in a new body that transformed into a blue van, Skids' first mission was to aid in preventing Shockwave from stealing sonic energy from a Brick Springstern concert, and he immediately took a liking to the music of Earth and the possibilities of what could be learned from interacting with mankind.
On the way to raid the Decepticon base, Skids accidentally clipped the car of hot-tempered Jake Dalrymple, who pursued the Autobots on their mission and distracted Skids just enough for Ravage to blast him off a cliff. Lying crippled and unable to transform, Skids was shortly discovered by register girl Charlene, who, thinking he was an ordinary van, had him fixed up with the intent of keeping him for herself. Skids was open to the idea at first, preferring to live a peaceful life to that of a warrior, but after another bout of road rage with Dalrymple, the unearthly stunts he performed left him no option but to reveal his true nature of Charlene. A somewhat unusual relationship ensued between the two, but more trouble soon came Skids's way in the form of Donny Finkleberg, who was being hunted down by Ravage. Fleeing from the Decepticon with Charlene and Donny, Skids hid out at an abandoned mining town, but yet again, the raging Dalrymple tracked him down and shattered his windshield, knocking him unconscious and into a dream where he died through his refusal to attack Megatron and save Charlene. Reawakening, he engaged Ravage in battle again, finally finding the strength to act, and causing his foe to tumble down a mineshaft. Skids then parted ways with Charlene, having come to the realisation that their two roles in life were too different for their relationship to work out.
Skids returned to the Ark with Finkleberg, and later embarked on a mission with him to track down a group of missing Autobots who had been captured by the Rapid Anti-robot Assault Team. Unfortunately, money still governed Finkleberg's actions, and he promptly sold out Skids to RAAT's leader Walter Barnett when Skids was deactivated for the night. Skids was disassembled for study, but when the Decepticons Runabout and Runamuck appeared on Earth and began defacing landmarks, RAAT operative Circuit Breaker was forced to enter into a deal with the Autobots to stop them. Skids and the other Autobots' components were combined to form a jury-rigged robot that Circuit Breaker used to stop the Battlechargers, but in return for their co-operation, Skids and the Autobots were allowed to go free.
On the return trip to the Ark, however, Skids was abruptly torn from reality when the future Decepticon Galvatron traveled back in time from 2006, shunting Skids into limbo to maintain temporal balance. Skids remained in limbo for two years while Galvatron ran amok on Earth, his mind steadily succumbing to the dream-illusions created by the monstrous creatures native to the realm.
With Galvatron's eventual destruction during the Time Wars, Skids was finally returned to Earth, but he had spent too long in Limbo -- the limbo-creatures had succeeded in shaping a reality from his dreams, and the nightmarish monsters that he had imagined materialised on Earth with him. Fleeing from the beasts, who desired to merge with Skids so that they might become more powerful, Skids lured them into a nuclear power plant which he had planned to detonate, wiping out the monsters and himself, but the remnants of the Autobot Wreckers -- joined by Catilla and Carnivac of the Decepticon Mayhem Attack Squad -- arrived to aid him, defeating the monsters. Skids joined with this group of Transformers, forming the Survivors.
Grimlock traveled to Hydrus Four to use Nucleon to revive the Dinobots. Returning to the Ark, he also pumped Nucleon into the med bays, reviving Skids and several others, and headed towards Cybertron. Grimlock used the Ark as a blunt instrument across Unicron's head, forcing Skids and the others to evacuate and join the battle against the Chaos Bringer.
When Carnivac ran off on his own to avenge Catilla's death, Skids and the other Survivors insisted Springer contact Grimlock and the Earthforce for back-up. Hooked up with these Autobot reinforcements, they stormed the Mayhem Attack Squad's tropical island base to save their friend. Skids personally confronted Octopunch in single combat, holding his own against the murderous Pretender.
On a later mission for Earthforce, Skids and the Survivors protected an oil tanker from Starscream, while the rest of the Decepticons were trying to distract the Autobots with an all-out assault on their base. Undeterred by the sleight-of-hand, the Survivors caught up with Starscream in the Gulf of Mexico. Skids used a flying leap by Carnivac as a distraction to catch Starscream with his liquid nitrogen cannon, and then Inferno knocked the Decepticon commander into the drink.
Like many of the Autobots who first appeared in 1985, the exact circumstances behind Skids joining the original group of Earth-based Autobots remain unknown. Skids first appeared as part of a strike force that was sent to stop the Insecticons from ravaging a human farm, only to meet with failure. Apparently injured in the battle, Skids underwent some repairs by Hoist upon return to Autobot Headquarters, where he responded incredulously to Bumblebee's report that he had retrieved robotic insecticide from the planet Floron 3 to deal with the Insecticon threat.
Later, Skids was part of a team sent to investigate bombs that were raining down on a city from a football stadium (the work of the Triple Changer Blitzwing). Blitzwing, however, had the stadium surrounded by a massive "Zone Defense" maze built by the Constructicons, and while attempting to traverse it, Skids crashed into Prowl and was driven over by tank-mode Blitzwing. Skids and the other Autobots' remains were then gathered by Scrapper, who built a throne for himself out of them. These Autobots were soon rescued and quickly rebuilt, and participated in the crises that ensued as Blitzwing, Astrotrain, Megatron and Devastator vied for Decepticon leadership. Following that skirmish, Prowl opined that Optimus Prime was the greatest leader in the universe, which prompted himself, Skids, Bluestreak and Tracks to form themselves into a throne for Prime; Prime refused to sit, saying that thrones were for Decepticons - he'd rather roll.
Skids was among the senior scientists (including Perceptor, Quickmix, and Rad) who questioned Rodimus Prime's plan to send out the best of the remaining Autobot science and research members to establish deep-space reconnaissance posts.
8.2 million years ago, Skids was in the Chamber of Ancients, observing the ceremony in which Optronix was given the Matrix of Leadership and transformed into Optimus Prime. Immediately, three Decepticon assassins attacked, and though Skids acted to protect the new Prime, his arm was cut off by Umbra.
Skids was repaired and he returned to his post at the Decagon. While Prime was absent, Skids picked up radar of incoming Decepticon invaders and a transmission from the force's field commander, Shockwave. Skids manned auto-defenses from inside the Decagon until they and their Omega Sentinel units were destroyed. Picking up their personal weaponry, Skids and the other Autobots were all that was left to defend Iacon, but defend it they did and Shockwave's forces were driven back.
During the Dark Ages, Skids served with Prowl's faction of Autobots. The team got involved in a skirmish with Trypticon and the giant Decepticon jumped onto a building in which Skids was trying to hide, crushing him. He survived though.
Skids was not among the Autobots who traveled to Earth aboard the Ark, remaining behind on Cybertron as the planet slipped into the deactivated state known as the Great Shutdown. When Shockwave began reactivating the planet and successfully unified the Autobot and Decepticon factions, Skids was a member of the council of Autobots who served to represent their faction in the alliance. Few of the council members were actually suited for leadership, however, and slowly found themselves giving more and more control to Shockwave. Eventually, the example shown by the Ark's returning crew and several homegrown Autobot insurgency cells tore the blinders from the council's optics, and Skids gathered his fellow councilmen in taking to the streets and ousting the Decepticons from power. Several council members would then accompany Prowl and the former Autobots of Earth back to that planet to construct Autobot City, and Skids may have been among them. Due to Dreamwave's self-destruction, we will never know.
Skids was the first of the Binaltech Autobots created simply with the intent of upgrading an existing, healthy Transformer, rather than in the name of repairing one that had been injured. Outfitted with self-repairing nanomechanical antibodies, Skids's body can, barring heavy damage in battle, theoretically function for thousands of years without maintenance. In both his original and new body, Skids dedicated time to studying the chronal warp on Dinobot Island, which gave rise to the Autobot strategy known as "Operation: Distant Thunder," a last-ditch scheme should the Decepticons tamper with the fabric of space-time. That horror occurred when Ravage used the Kronosphere to alter the timeline, and Skids prepared to initiate the plan and send Wheeljack's memory data back in time to avert the catastrophe.
The data probe was launched, but as the timeline was righting itself a Black Optimus Prime arrived, and manipulated the temporal wave to preserve the Binaltech timeline separate from the original continuity. Possessed by the Spark of the Protector, hailing from some future point in time, Black Optimus informed Skids and the others that their timeline was destined to produce great warriors capable of safeguarding the entire multiverse, and therefore must continue on its new course.